SUPERCLASS: GNATHOSTOMATA GEGENBAUR, 1874
= UNTERSTAMM EOTETRAPODA JAEKEL, 1911 (PISCES)

CLASS: ACANTHODII OWEN, 1846

= SUBCLASS: ACANTHODII OWEN, 1846
= SUBCLASS: ACANTHODIENS AGASSIZ, 1844
= SUBCLASS: ACANTHODEI OWEN, 1846
= SUBCLASS: ACANTHODII GIBEL, 1855
= SUBCLASS: ACANTHODINI LUTKNKE, 1868
= TRIBUS: ACANTHODIDES FRITSCH, 1893 (1895)
= SUBCLASS ACANTHODEA PARKER & HASWELL, 1897
= SUBCLASS ACANTHODIA DEAN, 1900
= SUBCLASS: ACANTHOESSI FOWLER, 1947

Order: INCERTAE SEDIS

Genus: Acanthospina VALUKEVICIUS, 2003
 A. irregulare VALUKEVICIUS, 2003 (Type)

Genus: Alienacanthus KULZCYCKI, 1957 (nomen dubium)
 A. malkowskii KULZCYCKI, 1957 (Type)

Genus: Antartonchus WHITE, 1968
A. glacilis WHITE, 1968 (Type)

Genus: Archaeacanthus KADE, 1858
 A. quadrisulcatus KADE, 1858

Genus: Asiacanthus LIU, 1948
 A. multituberculatus LIU, 1948 (Type)
 A. kao P’AN, 1964
A. suni P’AN, 1964

Genus: Byssacanthoides WOODWARD, 1921
 B. debenhami WOODWARD, 1921 (Type)

Genus: Campylodus ROHON, 1893
 C. sigmoides ROHON, 1893 (Type)
 C? delgadoi PRIEM, 1911

Genus: Cassidiceps GAGNIER & WILSON, 1996
 C. vermiculatus GAGNIER & WILSON, 1996 (Type)

Genus: Chelomodus ROHON, 1893 (nomen dubium)
 C. digitiferus ROHON, 1893 (Type)

Genus: Devononchus GROSS, 1940
 D. concinnus (GROSS, 1930) GROSS, 1940 (Type)
   = Onchus concinnus GROSS, 1930
D.  laevis (GROSS, 1933)
= Onchus laevis GROSS, 1933
D. tenuispinus (GROSS, 1933)
   = Onchus tenuispinatus GROSS, 1933
   = Archaeacanthus tenuispinatus (GROSS, 1933) NOVITSKAIA & OBRUCHEV, 1964
 D. ketleriensis GROSS, 1947

Genus: Ectopacanthus VALIUKEVICUS, 1979
 E. cristiformis VALKUKEVICUS, 1979 (Type)
 E? pusillus VALKUKEVICUS, 1998

Genus: Fungulodus
 F. rotundus

Genus: Gemuendolepis GROSS, 1973
 G. muelleri GROSS, 1973 (Type)

Genus: Granulacanthus HANKE, WILSON & LINDOE, 2001
 G. joenelsoni HANKE, WILSON & LINDOE, 2001 (Type)

Genus: Haplacanthus AGASSIZ, 1845
 H. marginalis AGASSIZ, 1845 (Type)
   = Onchus tenuisulacatus EICHWALD, 1846
H. ehrmanensis GROSS, 1940
H. perseenis GROSS, 1942
H. sp

Genus: Holmessla GUNNELL, 1931
 H. ornata GUNNELL, 1931 (Type)
 H. speni GUNNELL, 1933
 H. rhomboidalis GUNNELL, 1933

Genus: Homacanthus AGASSIZ, 1845
= Hoplonchus DAVIS, 1879
 H. gracilis (EICHWALD, 1844) AGASSIZ, 1845 (Type)
   = Hybodus gracilis EICHWALD, 1844
   = Homacanthus arcuatus AGASSIZ, 1844
 H. macrodus M’COY, 1848
   = Onchus subulatus AGASSIZ, 1843
H. microdus M’COY, 1848
 H. parvulus (NEWBERRY, 1875)
   = Ctenacanthus parvulus NEWBERRY, 1875
   = Hoplonchus parvulus NEWBERRY, 1889
 H. elegans (DAVIS, 1879)
   = Hoplonchus elegans DAVIS, 1879
 H. acinaciformis EASTMAN, 1903
 H. jaekeli (GROSS, 1933)
   = Ctenacanthus jaekeli GROSS, 1933
 H. sveteensis GROSS, 1942

Genus: Isodendracanthus VALIUKEVICUS, 1979
 I. ramiformis VALIUKEVICUS, 1979 (Type)

Genus: Lepracanthus OWEN, 1869 (nomen dubium)
 L. colei OWEN, 1869 (Type)

Genus: Lijiangichthys WANG, 2003
 L. lembodes WANG, 2003 (Type)

Genus: Maplemillia GROSS, 1973 (nomen dubium)
 M. costata GROSS, 1973 (Type)
M. sp

Genus: Monopleurodus PANDER, 1856
 M. ohhesaarensis PANDER, 1856 (Type)

Genus: Naulas AGASSIZ, 1845 (nomen dubium)
 N. sulcatus AGASSIZ, 1845 (Type)

Genus: Newsincanthus PAN & LIU, 1975
 N. planispinatus PAN & LIU, 1975 (Type)

Genus: Nodacosta GROSS, 1940
 N. pauli GROSS, 1940 (Type)
 N. gemuendensis (GROSS, 1933)
   = Ctenacanthus gemuendensis GROSS, 1933

Genus: Nodonchus WHITE, 1961
 N. bambusifer WHITE, 1961 (Type)
 N. rectus GAGNIER, TURNER, FRIMAN, SUAREZ-RIGLOS & JANVIER, 1988

Genus: Onychodus referred material:
 O. anglicus WOODWARD, 1888
 O. arcticus WOODWARD, 1889
 O. scoticus NEWTON, 1892
   = Protodus scoticus TRAQUAIR, 1898

Genus: Paucicanthus HANKE, 2002
 P. vanelsti HANKE, 2002 (Type)

Genus: Pinnacanthus BRYANT, 1934
 P. inequistriatus BRYANT, 1934 (Type)

Genus: Plectrodus AGASSIZ in MURCHISON, 1838
 P. pleiopristis AGASSIZ in MURCHISON, 1838 (Type)

Genus: Protodus WOODWARD, 1892
 P. jexi WOODWARD, 1892 (Type)
 P. sp
   = Stigmodus gracilis BROTZEN, 1934
   = Plectrodus hamatus BROTZEN, 1934
   = Dendrodus ariasigensis WHITEAVES, 1989

Genus: Rabdacanthus PANDER, 1856 (nomen dubium)
  R. truncatus PANDER, 1856 (Type)
R. sp

Genus: Sentacanthus KULCZYCKI, 1957 (nomen dubium)
 S. zelichowskae KULCZYCKI, 1957 (Type)

Genus: Watsonacanthus VALIUKEVICUS, 1979
 W. oervigi VALIUKEVICUS, 1979 (Type)
 W. costatus VALUIKEVICIUS, 2003
W. sp

Genus: Yealepis BURROW & YOUNG, 1999
 Y. douglasi BURROW & YOUNG, 1999 (Type)

Genus: Yunnanacanthus LU, 1973
 Y. cuifengshanensis LU, 1973 (Type)

Family: SINACANTHIDAE Zhu, 1998

Genus: Sinacanthus P’AN, 1957
 S. micracanthus (CHAPMAN, 1917) ZHU, 1998
  = Physonemus. micracanthus CHAPMAN, 1917
S. wuchangensis P’AN, 1957 (Type)
 S. fanchunensis LIU, 1973
 S. triangulatus PAN & LIU, 1975
 S. boliviensis GAGNIER, TURNER, FRIMAN, SUAREZ-RIGLOS & JANVIER, 1988
S. sp

Family: MACHAERACANTHIDAE Burrow & Young, 2005

Genus: Machaeracanthus NEWBERRY, 1857
= Dinacanthodes FRITSCH, 1893 (1895)
= Machaerius ROUAULT, 1858
 M. peracutus NEWBERRY, 1857 (Type)
 M. major NEWBERRY, 1857
 M. peracutus NEWBERRY, 1857
 M. sulcatus NEWBERRY, 1857
 M. archiaci (ROUAULT, 1858)
   = Machaerus archiaci ROUAULT, 1858
 M. larteti (ROUAULT, 1858)
   = Machaerius larteti ROUAULT, 1858
M. bohemicus (BARRANDE, 1872)
   = Ctenacanthus bohemicus BARRANDE, 1872
   = Dinacanthodes sussii FRITSCH 1893 (1895)
   = Cheiracanthoides stonehouensis LEGAULT, 1968
 M. polonicus GURICH, 1901
 M. longaevus EASTMAN, 1907
 M. kayseri KEGEL, 1913
  = Machaeracanthus westfalicus PFEIFFER, 1938
 M. minor BRYANT, 1934
 M. retusus WELLS, 1940
 M. pectinatus BURROW & YOUNG, 2005
 M. hunsruekianum SUKKAMP & BURROW, 2007
M. sp

ORDER: TCHUNACANTHIDAE KARATAJUTE-TALIMAA & SMITH, 2003 (FOR 2002)

Family: LENACANTHIDAE Karatajute-Talimaa & Smith, 2003 (FOR 2002)

Genus: Lenacanthus KARATAJUT-TALIMAA & SMITH, 2003 (For 2002)
 L. priscus KARATAJUT-TALIMAA & SMITH, 2003 (For 2002) (Type)

Family: TCHUNACANTHIDAE karatajute-Talimaa & Smith, 2003 (for 2002)

Genus: Fecundosquama VALIUKEVICUS, 2004
 F. basiglobosa VALIUKEVICUS, 2004 (Type)

Genus: Tchunacanthus KARATAJUT-TALIMAA & SMITH, 2003 (For 2002)
 T. obruchevi KARATAJUT-TALIMAA & SMITH, 2003 (For 2002) (Type)

ORDER: ISCHNACANTHIFORMES BERG, 1940
= ORDER: ISCHNACANTHIDA BERG, 1940
= ORDER: ISCHNACANTHOMORPHI ORVIG, 1957
= ORDER: ISCHNACANTHIDA NOVITSKAIA & OBRUCHEV, 1964

Incertae sedis

Genus: Serradentus NEWMAN, BURROW & den BLAAUWEN, 2020
S. armstrongi NEWMAN, BURROW & den BLAAUWEN, 2020 (Type)
S. sp

Family: PODOLICANTHIDAE Voichyshyn & Szaniawski, 2018

Genus: Drygantacanthus VOICHYSHYN & SZANIAWSKI, 2018
D. semirotunda VOICHYSHYN & SZANIAWSKI, 2018 (Type)

Genus: Kasperacanthus VOICHYSHYN & SZANIAWSKI, 2018
K. serratus VOICHYSHYN & SZANIAWSKI, 2018 (Type)

Genus: Podolicanthus VOICHYSHYN & SZANIAWSKI, 2012
P. zychi VOICHYSHYN & SZANIAWSKI, 2012 (Type)
P. sp

Gen. sp indet.

Family: ISCHNACANTHIDAE Woodward, 1891
= Family: ONCHIDAE Jordan, 1923
= Family: PROTODONTIDAE Woodward in Zittel, 1932

Genus: Acanthodopsis HANCOCK & ATTHEY, 1868
A. wardi HANCOCK & ATTHEY, 1868 (Type)
= Acanthodopsis egertoni HACOCK & ATTHEY, 1868
A. microdon TRAQUAIR, 1894
A. sp

Genus: Apateacanthus WOODWARD, 1891
 A. vetustus (CLARKE, 1885) WOODWARD, 1891 (Type)
   = Pristacanthus vetustus CLARKE, 1885
 A. peculiaris HUSSAKOF, 1913

Genus: Arenaceacanthus VALIUKEVICIUS, 2004
 A. arcuatacanalis VALIUKEVICIUS, 2004 (Type)

Genus: Atopacanthus HUSSAKOF & BRYANT, 1918
 A. dentatus HUSSAKOF & BRYANT, 1918 (Type)
 A. peculiaris (HUSSAKOF, 1913)
   = Apatacanthus peculiaris HUSSAKOF, 1913
 A? ambrockensis OTTO, 1999
A. sp

Genus: Bracteatacanthus VALIUKEVICIUS, 2004
 B. assiduous VALUKEVICIUS, 2004 (Type)

Genus: Cambaracanthus LINDLEY, 2000
 C. goodhopensis LINDLEY, 2000 (Type)

Genus: Cavanacanthus LINDLEY, 2000
 C. warrooensis LINDLEY, 2000 (Type)

Genus: Doliodus TRAQUAIR, 1893
 D.  problematicus (WOODWARD, 1892) TRAQUAIR, 1893 (Type)
   = Diplodus problematicus WOODWARD, 1892
   = Dissodus problematicus MILLER 1892

Genus: Erymnacanthus BLAIS, HERMUS & WILSON, 2015
E. clivus BLAIS, HERMUS & WILSON, 2015 (Type)

Genus: Euryacanthus BLAIS, HERMUS & WILSON, 2015
E. rugosus BLAIS, HERMUS & WILSON, 2015 (Type)

Genus: Gansuichthys WANG, WANG, ZHANG & WANG, 1998
 G. liui WANG, WANG, ZHANG & WANG, 1998 (Type)

Genus: Garralepis BURROW, 2002
G. simplex BURROW, 2002 (Type)
G. sp

Genus: Gomphonchus GROSS, 1971
 G. sandelensis (PANDER, 1856) GROSS, 1971 (Type)
   = Thelolepis volborthi ROHON, 1893
   = Onchus roemeri HOPPE 1931
   = Gomphodus longifromis BROTZEN, 1934
   = Gomphodus rhomboidalis BROTZEN, 1934
   = Acanthodes devonicus BROTZEN, 1934
   = Acanthodes laevigatus BROTZEN, 1934
   = Acanthoides inornatus BROTZEN, 1934
   = Acanthoides exscuplatus BROTZEN, 1934
   = Acanthoides undulatus BROTZEN, 1934
   = Poracanthodes punctatus BROTZEN, 1934
   = Poracanthodes asper BROTZEN, 1934
   = Poracanthodes biblicus LEHMAN, 1937
   = Gomphodus hoppei GROSS, 1968
 G. globosus (HOPPE 1931) (nomen dubium)
 G. curvotriangulatus (LEHMAN, 1937) (nomen dubium)
 G. sandalensis (GROSS, 1947)
   = Gomphodus sandalensis GORSS, 1947
 G. pluriformis GAGNIER, TURNER, FRIMAN, SUAREZ-RIGLOS & JANVIER, 1988
G. bogongensis BURROW, 1997
 G. tauragensis VALIUKEVICIUS, 1998
 G. medicostatus VERGOOSSEN, 1999
 G. boekschoteni Vergossen, 1999
 G. fromensis BURROW, 2002
 G.nordicus VALIUKEVICIUS, 2003
 G? minutus VALIUKEVICIUS, 2004
G. sp

Genus: Grenfellacanthus LONG, BURROW & RITCHIE, 2004
 G. zerinae LONG, BURROW & RITCHIE, 2004 (Type)

Genus: Helenacanthus BRYANT, 1934
  H. incurvus BRYANT, 1934 (Type)

Genus: Ischnacanthus POWRIE, 1864
= Ictinocephalus PAGE, 1859 (nomen nudum)
 I. gracilis (EGERTON, 1861) POWRIE, 1864 (Type)
   = Diplacanthus gracilis EGERTON, 1861
= Onychodus scoticus E. T. NEWTON, 1892
I? anglicus WHITE, 1961
I. kingi WHITE, 1961
I. wickhami WHITE, 1961
I? scheii SPJELDNAES, 1967

Genus: Lietuvacanthus VALIUKEVICIUS, 1998
 L. fossulatus VALIUKEVICIUS, 1998 (Type)

Genus: Marsdenius WELBURN, 1902
 M. summiti WELBURN, 1902 (Type)
 M. acuta WELBURN, 1902

Genus : Nerepisacanthus BURROW & RUDKIN, 2014
Nerepisacanthus denisoni BURROW & RUDKIN, 2014 (Type)

Genus: Persacanthus JANVIER, 1977
 P. kermanensis JANVIER, 1977 (Type)
 P. simpsonensis REED, 1986
P. sp

Genus: Plectrodus AGASSIZ in MURCHISON, 1839
 P. mirabilis AGASSIZ in MURCHISON, 1839 (Type)

Genus: Rockycampacanthus LONG, 1986a
 R. milesi LONG, 1986a (Type)

Genus: Rohonilepis VALIUKEVICIUS, 2004
 R. breviornatus VALIUKEVICIUS, 2004 (Type)

Genus: Taemasacanthus LONG, 1986a
 T. erroli LONG, 1986a (Type)
 T. porca LINDLEY, 2000
 T. narrengullensis LINDLEY, 2001
 T. cooradigbeensis LINDLEY, 2001

Genus: Tricuspicanthus BLAIS, HERMUS & WILSON, 2015
T. gannitus BLAIS, HERMUS & WILSON, 2015 (Type)
T. pisculus BLAIS, HERMUS & WILSON, 2015

Genus: Trundlelpis BURROW, 1997
 T. cervicostulata BURROW, 1997 (Type)

Genus: Uraniacanthus MILES, 1973
U. spinosus MILES, 1973 (Type)

Genus: Xylacanthus ORVIG, 1967
 X. grandis ORVIG, 1967 (Type)
X. minutus GAGNIER & GOUJET, 1997
X. kenstewarti HANKE, WILSON & LINDOE, 2001

Genus: Youngacanthus WANG, 1984
 Y. gracilis WANG, 1984 (Type)

Family: GLADOBRANCHIDAE Bernacsek, & Dineley, 1997

Genus: Gladiobranchus BERNACSEK, & DINELEY, 1977
 G. probaton BERNACSEK, & DINELEY, 1977 (Type)

Family: PORACANTHODIDAE Vergoossen, 1997

Genus: Acanthopora VALUKEVICIUS, 2003
 A. transitans VALUKEVICIUS, 2003 (Type)

Genus: Articacanthus VALUKEVICIUS, 2003
 A. bicostatus VALUKEVICIUS, 2003 (Type)

Genus: Gomphonchoporus VERGOOSSEN, 1999
 G. hoppei (GROSS, 1947) VERGOOSSEN, 1999 (Type)  
   = Gomphodus hoppei GROSS, 1947
   = Ghomphodus irregularis LEHMAN, 1937

Genus: Poracanthodes BROTZEN, 1934
= Brotzenlepis VERGOOSSEN, 1997
 P. punctatus BROTZEN, 1934 (Type)
 P. menneri VALIUKEVICIUS, 1992
 P. zoigenensis WANG, WANG, ZHANG & WANG, 1998
 P.subporosus VALIUKEVICIUS, 1998
 P? lehmani VERGOOSSEN, 1999
P. canadensis BURROW, VERGOOSSEN, TURNER, UYENEO & THORSTEINSSON, 1999
P. sulcatus VALIUKEVICIUS, 2004
P. marssae VALIUKEVICIUS, 2004
P. sp

Genus: Radioporacanthodes VERGOOSSEN, 1999
 R. porosus (BROTZEN, 1934) VERGOOSEEN, 1999 (Type)
   = Poracanthodes porosus BROTZEN, 1934
 R. qujingensis (WANG & DONG, 1989)
   = Poracanthodies qujiangensis WANG & DONG 1989
R. sp

Genus: Zemlyacanthus VERGOOSSEN, 1997

 ORDER: CLIMATIIFORMES BERG, 1940
= ORDER: CLIMATIIDA BERG, 1940
= ORDER: CLIMATIOMORPHI ORVIG, 1957
= ORDER: CLIMATIIDA FOWLER, 1965

Incertae sedis

Genus: Nobilesquama VALIUKEVICIUS & BURROW, 2005
N. longipostera (VALIUKEVICIUS, 2003) VALIUKEVICIUS & BURROW, 2005 (Type)
N. minilonga (VALIUKEVICIUS, 2003) VALIUKEVICIUS & BURROW, 2005
N. watsoni (VALIUKEVICIUS, 2003) VALIUKEVICIUS & BURROW, 2005
= Nostolepis watsoni VALIUKEVICIUS, 2003

Genus: Nostovicina VALIUKEVICIUS & BURROW, 2005
N. applicata (VIETH, 1980) VALIUKEVICIUS & BURROW, 2005
= Nostolepis applicata VIETH, 1980
N. multicostata (VIETH, 1980) VALIUKEVICIUS & BURROW, 2005
= Nostolepis multicostata VIETH, 1980
N. guangxiensis (WANG, 1992) VALIUKEVICIUS & BURROW, 2005
= Nostolepis guangxiensis WANG, 1992
 = “Nostolepisguangxiensis BURROW, LONG & TURNER, 1998
N.  lacrima (VALIUKEVICIUS, 1994) VALIUKEVICIUS & BURROW, 2005
= Nostolepis lacrima VALIUKEVICIUS, 1994
N. laticristata (VALIUKEVICIUS, 1994) VALIUKEVICIUS & BURROW, 2005
= Nostolepis laticristata VALIUKEVICIUS, 1994
athleta (VALIUKEVICIUS, 1994) VALIUKEVICIUS & BURROW, 2005
= Nostolepis athleta VALIUKEVICIUS, 1994
N. curiosa (VALIUKEVICIUS, 1994) VALIUKEVICIUS & BURROW, 2005
= Nostolepis curiosa VALIUKEVICIUS, 1994
N. multangula (VALIUKEVICIUS, 1994) VALIUKEVICIUS & BURROW, 2005
= Nostolepis multangula VALIUKEVICIUS, 1994
N. tareyensis (VALIUKEVICIUS, 1994) VALIUKEVICIUS & BURROW, 2005
= Nostolepis tareyensis VALIUKEVICIUS, 1994
N. spina (VALIUKEVICIUS, 1994) VALIUKEVICIUS & BURROW, 2005
= Nostolepis spina VALIUKEVICIUS, 1994
N. halli (BLOM, 1999) VALIUKEVICIUS & BURROW, 2005
N. fragila (VALIUKEVICIUS, 2003) VALIUKEVICIUS & BURROW, 2005 (Type)
N. paravolborthi (VALIUKEVICIUS, 2003) VALIUKEVICIUS & BURROW, 2005
N. platycrista (VALIUKEVICIUS, 2003) VALIUKEVICIUS & BURROW, 2005
N. timanica (VALIUKEVICIUS, 2003) VALIUKEVICIUS & BURROW, 2005
= Nostolepis timanica VALIUKEVICIUS, 2003

Genus: Peregrinosquama VALIUKEVICIUS & BURROW, 2005
P. costata (VALIUKEVICIUS, 2003) VALIUKEVICIUS & BURROW, 2005 (Type)
P. oervigi (SOBOLEV, KARATAJUTE-TALIMAA & VALIUKEVICIUS, 1988) VALIUKEVICIUS & BURROW, 2005
P. sp

SUBORDER: CLIMATIFORMOIDEI MILES, 1966

Family: CLIMATIIDAE Berg, 1940
= Family: BRACHYACANTHIDAE Moy-Thomas, 1939
= Family: BROCHADMONIDAE Bernacsek & Dineley, 1977

Genus: Brachyacanthus EGERTON, 1860
 B. scutiger EGERTON, 1860 (Type)

Genus: Brochoadmones BERNACESK & DINELEY, 1977
 B. milesi BERNACESK & DINELEY, 1977 (Type)

Genus: Canadalepis VIETH, 1980
 C. linguiformis VIETH, 1980 (Type)
C. sp

Genus: Cheiracanthoides WELLS, 1944
= Helolepis WELLS, 1944
 C. comptus WELLS, 1944 (Type)
   = Cheiracanthoides breviplicatus WELLS, 1944
   = Cheiracanthoides comis WELLS, 1944
   = Cheiracanthoides venustus WELLS, 1944
   = Helolepis bellarugosus WELLS, 1944
  = Helolepis venustuloides WELLS, 1944
 C. rarus VALIUKEVICIUS, 1994
 C. planus VALIUKEVICIUS, 1998
 C. nativus VALIUKEVICIUS, 1998
 C. estonicus VALIUKEVICIUS, 1998
C. sp

Genus: Climatius AGASSIZ, 1845
 C? ornatus (AGASSIZ, 1837)
   = Ctenacanthus ornatus AGASSIZ, 1837
 C. reticulatus AGASSIZ, 1845 (1844)
 C. latispinosus (WHITEAVES, 1881)
   = Homacanthus gracilis WHITEAVES, 1889
 C. enodiscota GAGNIER, TURNER, FRIMAN, SUAREZ-RIGLOS & JANVIER, 1988
 C. sp

Genus: Endemolepis VALIUKEVICIUS, 1998
 E. inconstans VALIUKEVICIUS, 1998 (Type)

Genus: Erriwacanthus ORVIG, 1967
 E. falcatus ORVIG, 1967 (Type)
 E. manbrookensis MILES, 1973
E. sp

Genus: Iranolepis HAIRAPETIAN, VALIUKEVICIUS & BURROW, 2006
I. ginteri HAIRAPETIAN, VALIUKEVICIUS & BURROW, 2006 (Type)

Genus: Lalicanthus KARTAJUTE-TALIMAA, 1986
 L. singularis KARTAJUTE-TALIMAA, 1986 (Type)
 L. ligeriensis VIDAL, GOUJET, JANVIER & LARDEUX, 1994

Genus: Latviacanthus SCHULTZE, 1982
 L. ventspilsensis SCHULTZE, 1982 (Type)

Genus: Lupopsyrus BERNACSEK & DINELEY, 1977
 L. pygmaeus DINELEY, 1977 (Type)

Genus: Nostolepis PANDER, 1856
= Acantholepis BROTZEN, 1934, non NEWBERRY, 1875
= Belacanthus BROTZEN, 1934
= Conolepis BROTZEN, 1934
= Dendracanthus LEHMAN, 1937
= Diplacanthoides BROTZEN, 1934
= Dontancathus LEHMAN, 1937
= Rhabdidodus ROHON, 1893
N. striata PANDER, 1856 (Type)
   = Onchus curvatus PANDER, 1856
   = Onchus granulatus ROEMER, 1885
   = Diplacanthoides balticus BROTZEN, 1934
  = Nostolepis balticus (BROTZEN, 1934)
  = Diplacanthoides rugosus BROTZEN, 1934
  = Nostolepis rugosus (BROTZEN, 1934)
  = Diplacanthoides furcatus BROTZEN, 1934
  = Nostolepis furcatus (BROTZEN, 1934)
  = Diplacanthoides insignis BROTZEN, 1934
  = Nostolepis insignis (BROTZEN, 1934)
  = Diplacanthoides elegans BROTZEN, 1934
  = Nostolepis elegans (BROTZEN, 1934)
  = Diplacanthoides crassus BROTZEN, 1934
  = Nostolepis crassus (BROTZEN, 1934)
   = Diplacanthoides bisulcatus BROTZEN, 1934
  = Nostolepis bisulcatus (BROTZEN, 1934)
   = Diplacanthoides globosus BROTZEN, 1934
  = Nostolepis globosus (BROTZEN, 1934)
   = Diplacanthoides hoppei BROTZEN, 1934
  = Nostolepis hoppei (BROTZEN, 1934)
   = Conolepis conulus BROTZEN, 1934
  = Nostolepis conulus (BROTZEN, 1934)
   = Conolepis multicornis BROTZEN, 1934
  = Nostolepis multicornis (BROTZEN, 1934)
   = Gomphodus trinucleatus BROTZEN, 1934
  = Nostolepis trinucleatus (BROTZEN, 1934)
   = Gomphodus magnisquamosus BROTZEN, 1934
  = Nostolepis magnisquamosus (BROTZEN, 1934)
   = Gomphodus ornatus BROTZEN, 1934
  = Nostolepis ornatus (BROTZEN, 1934)
   = Cheiracanthus vogti BROTZEN, 1934
  = Nostolepis vogti (BROTZEN, 1934)
  = Belacanthus carinatus BROTZEN, 1934
  = Nostolepis carinatus (BROTZEN, 1934)
   = Dendracanthus ? cristatus LEHMAN, 1937
    = Nostolepis? cristatus (LEHMAN, 1937)
   = Dendracanthus fibrulatus LEHMAN, 1937
    = Nostolepis fibrulatus (LEHMAN, 1937)
  = Dendracanthus radiocostatus LEHMAN, 1937
  = Nostolepis radiocostatus (LEHMAN, 1937)
   = Dendracanthus stensioi LEHMAN, 1937
  = Nostolepis stensioi (LEHMAN, 1937)
   = Diplacanthoides compressus LEHMAN, 1937
   = Nostolepis compressus (LEHMAN, 1937)
   = Diplacanthoides decoratus LEHMAN, 1937
  = Nostolepis decoratus (LEHMAN, 1937)
   = Diplacanthoides dilobatus LEHMAN, 1937
   = Nostolepis dilobatus  (LEHMAN, 1937)
   = Nostolepis multicostatus (LEHMAN, 1937)
   = Diplacanthoides multicostatus LEHMAN, 1937
   = Nostolepis multilobatus (LEHMAN, 1937)
   = Diplacanthoides multilobatus LEHMAN, 1937
   = Nostolepis parvolobatus (LEHMAN, 1937)
   = Diplacanthoides parvolobatus LEHMAN, 1937
   = Diplacanthoides retroincisuratus LEHMAN, 1937
   = Nostolepis retronicisuratus (LEHMAN, 1937)
   = Diplacanthoides sinuosus LEHMAN, 1937
   = Nostolepis sinuosus (LEHMAN, 1937)
   = Diplacanthoides trilobatus LEHMAN, 1937
   = Nostolepis trilobatus (LEHMAN, 1937)
   = Diplacanthoides unguiparatus LEHMAN, 1937
   = Nostolepi. unguiparatus (LEHMAN, 1937)
   = Dontacanthus costatus LEHMAN, 1937
   = Nostolepis costatus (LEHMAN, 1937)
   = Gomphodus foliosus LEHMAN, 1937
   = Nostolepis foliosus (LEHMAN, 1937)
   = Gomphodus inaequalobatus LEHMAN, 1937
   = Nostolepis inaequalobatus (LEHMAN, 1937)
   = Diplacanthoides irregularis LEHMAN, 1937
   = Nostolepis irregularis (LEHMAN, 1937)
   = Diplacanthoides laterolobatus LEHMAN, 1937
   = Nostolepis laterolobatus (LEHMAN, 1937)
   = Diplacanthoides multiincisuratus LEHMAN, 1937
   = Nostolepis multiincisuratus (LEHMAN, 1937)
   = Diplacanthoides trilingulatus LEHMAN, 1937
   = Nostolepis trilingulatus (LEHMAN, 1937)
N. parvidens (ROHON, 1893)
   = Rhabdiodus parvidens ROHON, 1893
N? dentoides HOPPE, 1931
N? lata HOPPE, 1931
N? parva HOPPE, 1931
N? pseudocostata HOPPE, 1931
N. robusta (BROTZEN, 1934)
    = Diplacanthoides robusta BROTZEN, 1934
N. incurvus (BROTZEN, 1934)
    = Acanthoides incurvus BROTZEN, 1934
N. coronata (BROTZEN, 1934)
   = Acantholepis coronata BROTZEN, 1934
N. typicus (BROTZEN, 1934)
   = Belacanthus typicus BROTZEN, 1934
N. huckei (BROTZEN, 1934)
   = Diplacanthoides huckei BROTZEN, 1934
N. robustus (BROTZEN, 1934)
   = Diplacanthoides robustus BROTZEN, 1934
N. alatus (BROTZEN, 1934)
   = Gomphodus alatus BROTZEN, 1934
N. erraticus (BROTZEN, 1934)
   = Poracanthoides erraticus BROTZEN, 1934
N. gracilis GROSS, 1947
N. costata GOUJET, 1976
N. arctica VIETH, 1980
N. striata TURNER & PICKETT, 1982
N. kernavensis VALIUKEICIUS, 1985
N. minima VALIUKEVICIUS, 1994
N. tewonensis WANG, WANG, ZHANG & WANG, 1998
N. gaujensis VALIUKEVICIUS, 1998
N. linleyensis MILLER & MARSS, 1999
N. mingyinensis WANG, 2003
N. decora VALIUKEVICIUS, 2003
N. fragilis VALIUKEVICIUS, 2003
N.parathelta VALIUKEVICIUS, 2003
N.kozhymica VALIUKEVICIUS, 2003
N.terraborea VALIUKEVICIUS, 2003
N.amplifica VALIUKEVICIUS, 2003
N.magnicostata VALIUKEVICIUS, 2003
N.consueta VALIUKEVICIUS, 2003
N.musca VALIUKEVICIUS, 2003
N. alifera VALIUKEVICIUS, 2004
N. latvica VALIUKEVICIUS, 2004
N. qujingensis LI, CUI, ANDREEV, SHAO, WANG, PENG & ZHU, 2021
N. digitus LI, CUI, ANDREEV, SHAO, WANG, PENG & ZHU, 2021
N. sp

Genus: Nostolepoides BURROW, 1997
 N. platymarginata BURROW, 1997 (Type)

Genus: Onchus AGASSIZ, 1837 (1843)
 O. murchisoni AGASSIZ, 1837 (1843) (Type)
 O. tenuistriatus AGASSIZ, 1837 (1843)
 O. falcatus AGASSIZ, 1843 (nomen nudum)
 O. sublaevis AGASSIZ, 1845
 O. heterogyrus AGASSIZ, 1845 (1844) (nomen dubium)
   = Heteracanthus heterogyrus (AGASSIZ, 1845 (1844)) WOODWARD, 1891
   = Gamphacanthus heterogyrus BRYANT, 1929
 O. decorus PHILLIPS, 1848 (nomen dubium)
 O. dubius PANDER, 1856 (nomen dubium)
 O. tricarinatus PANDER, 1856 (nomen dubium)
 O. compressus EICHWALD, 1860 (nomen dubium)
 O. major ETHERIDEGE in SYMONDS, 1872
 O. clintoni CLAYPOLE, 1885
 O. rectus EASTMAN, 1899 (nomen nudum)
 O. graptolitarum FRITSCH, 1907
   = Onchus graptolithorum GROSS, 1950 (sic)
 O. siluricus FRITSCH, 1907
   = Onchus silesiacus GROSS, 1950 (sic)
 O. latus LERICH, 1931
 O. penetrans BRYANT, 1932
 O. peracutus BRYANT, 1934
 O. overathensis GROSS, 1937
 O? brotzeni LEHMAN, 1937 (nomen dubium)
 O. verus SCHMIDT, 1954
 O. rarus LIEPIN’SH, 1959
 O. wheathillensis WHITE, 1961
 O? besomeneis WHITE, 1961 (nomen dubium)
 O. seriponensis GAGNIER, TURNER, FRIMAN, SUAREZ-RIGLOS & JANVIER, 1988
 O. sicaeformis GAGNIER, TURNER, FRIMAN, SUAREZ-RIGLOS & JANVIER, 1988
O. sp

Genus: Parexus AGASSIZ, 1845
 P. recurvus AGASSIZ, 1845 (Type)
 P. falcatus POWRIE, 1870
 P. sp
   = Farnellia tuberculata TRAQUIAR, 1898

Genus: Pruemolepis VIETH-SCHREINER, 1983
 P. wellsi VIETH-SCHREINER, 1983 (Type)

Genus: Ptomacanthus MILES, 1973
 P. anglicus MILES, 1973 (Type)

Genus: Sabrinacanthus MILES, 1973
 S. arcuatus (AGASSIZ, 1837) MILES, 1973 (Type)
   = Onchus arcuatus AGASSIZ, 1837

Genus: Tareyacanthus VALIUKEVICIUS, 1994
T. magnificus VALILUKEVICIUS, 1994 (Type)
 T. dissectus VALIUKEVICIUS, 1994

Genus: Wetteldorfia VIETH-SCHREINER, 1983
W. triangula VIETH-SCHREINER, 1983 (Type)
W. sp

Family: EUTHACANTHIDAE Berg, 1940

Genus: Euthacanthus POWRIE, 1864
 E. macnicoli POWRIE, 1864 (Type)
 E. elegans POWRIE, 1870
 E. gracilis POWRIE, 1870
 E. grandis POWRIE, 1870
 E. curtus POWRIE, 1870

Family: VESPERALIIDAE Valiukevicius & Burrow, 2005

Genus: Acanthacanthus VALIUKEVICUS, 2003
 A. ornatus VALIUKEVICUS, 2003 (Type)

Genus: Vesperalia VALIUKEVICUS, 2004
 V. perplexa VALIUKEVICUS, 2004 (Type)

Family: ACRITOLEPIDAE Valiukevicius & Burrow, 2005

Genus: Acritolepis VALIUKEVICIUS, 2003
 A. ushakovi VALIUKEVICIUS, 2003 (Type)
 A. urvantsevi VALIUKEVICIUS, 2003

Genus: Monospina VALIUKEVICUS, 2003
M. erecta VALIUKEVICIUS, 2003 (Type)

Genus: Pechoralepis VALIUKEVICIUS & BURROW, 2005
 P. zinaidae (VALIUKEVICIUS, 2003) VALIUKEVICIUS & BURROW, 2005 (Type)
 P. valentinae (VALIUKEVICIUS, 2003) VALIUKEVICIUS & BURROW, 2005
 P. advensis (VALIUKEVICIUS, 2003) VALIUKEVICIUS & BURROW, 2005
P. juozasi  BURROW, LONG & TRINAJSTIC, 2009

ORDER: DIPLACANTHIFORMES, BERG, 1940
= SUBORDER: DIPLACANTHIDEI MILES, 1966

Family: DIPLACANTHIDAE Woodward, 1891
= Family: DIPLOCANTHIDAE Fowler, 1947 (sic)

Genus: Diplacanthus AGASSIZ, 1844
= Diplocanthus DUFF 1842, incorrectly transcribed from AGASSIZ MS.
= Rhadinacanthus TRAQUAIR, 1888
 D. longispinius AGASSIZ, 1844 (Type)
= Rhadinacanthus longispinius (AGASSIZ, 1844) TRAQUAIR, 1888
D. crassisimus DUFF 1842
  = Diplocanthus striatus
  = Diplocanthus striatulus
   = Diplocanthus crassipinus AGASSIZ, 1844
   = Diplocanthus gibbus M’COY, 1848
D. horridus WOODWARD, 1892
   = Rhadinacanthus horridus (WOODWARD, 1892) TRAQUAIR, 1888
D. tenuistratus TRAQUAIR, 1894
   = Homacanthus borealis TRAQUAIR, 1893
D? carinatus GROSS, 1973
D. gravis VALIUKEVICIUS, 1988
D. ellsi GAGNIER, 1996
   = Diplacanthus crassimus HAY, 1902
   = Diplacanthus stiatus EASTMAN, 1907
   = Diplacanthus horridus RUSSELL, 1951 non WOODWARD, 1892
 D. acus GESS, 2001
 D. poltnigi VALUIKEVICIUS, 2003
 D. solidus VALUIKEVICIUS, 2003

Genus: Florestacanthus BURROW, JANVIER & VILLARROEL, 2003
 F. morenoi BURROW, JANVIER & VILLARROEL, 2003 (Type)

Genus: Gladiobranchus BERNACSEK & DINELEY, 1977
 G. probaton BERNACSEK & DINELEY, 1977 (Type)

Genus: Jaufolepis FOREY, YOUNG, & McCLURE, 1992
 J. striata FOREY, YOUNG & McCLURE, 1992 (Type)

Genus: Milesacanthus YOUNG & BURROW, 2004
M. antarctica YOUNG & BURROW, 2004 (Type)
M. ancestralis BURROW, LELIEVRE & JANJOU, 2006

Genus: Ptychodictyon GROSS, 1973
P. sulcatum GROSS, 1973 (Type)
P. rimosum GROSS, 1973
P. distinctum VALIUKEVICIUS, 1979

Genus: Rhadinacanthus TRAQUAIR, 1888
 R. longispinus (AGASSIZ 1844) TRAQUAIR, 1888
   = Rhadinacanthus TRAQUAIR, 1888  (Type genus)
   = Diplacanthus longispinus AGASSIZ 1844
   = Diplocanthus perarmatus M’COY, 1848
R? balticus GROSS, 1973
R. primaris VALIUKEVICIUS, 1986
R. multisculcatus VALIUKEVICICUS, 1988
R. deltosquamosus PINAKINA & MARSS, 2018
R. sp

Gen. sp indet.

Family: TETANOPSYRIDAE Hanke, DAVIS & WILSON, 2001

Genus: Tetanopsyrus GANGIER, HANKE & WILSON, 2001
 T. lindoei GANGIER, HANKE & WILSON, 2001 (Type)
 T. breviacanthias HANKE, DAVIS & WILSON, 2001

Family: CULMACANTHIDAE

Genus: Culmacanthus LONG, 1983
 C. stewarti LONG, 1983 (Type)
 C. antartica YOUNG, 1989
 C. pambulensis YOUNG, 1989

Family: GYRACANTHIDAE Woodward, 1906
= ORDER: GYRACANTHIFORMES BERG, 1940
= ORDER: GYRACANTHIDA NOVITSKAIA & OBRUCHEV, 1964
= Family: GYRACANTHIDIDAE Miles, 1971

Genus: Aganacanthus TRAQUAIR, 1884
 A. striatulus TRAQUAIR, 1884 (Type)

Genus: Antacanthus DEWALQUE, 1877
 A. insignis DEWALQUE, 1877 (Type)

Genus: Gyracanthus AGASSIZ, 1837 (1843)
 G. formosus AGASSIZ 1837 (1843) (Type)
   = Gyracanthus tuberculatus AGASSIZ 1837 (1843)
 G. alnwicensis AGASSIZ 1837 (1843)
 G. obliquus M’COY 1848
G. quadricornis (OWEN, 1867) (nomen dubium)
   = Mitroduos quadricornis OWEN, 1867 (nomen dubium)
 G. duplicatus DAWSON, 1868
 G. magnificus DAWSON, 1868
 G. alleni NEWBERRY, 1873
 G. compressus NEWBERRY, 1873
 G. nobilis TRAQUAIR, 1883
 G. youngii TRAQUAIR, 1883
 G? cordatus St. JOHN & WORTHEN, 1883 (nomen dubium)
 G. inoratus NEWBERRY, 1889
 G. sherwoodi NEWBERRY, 1889
 G. incurvus TRAQUAIR, 1890
 G. rectus TRAQUAIR, 1890
 G. faliciformis TRAQUAIR, 1902
 G. murrayi (McCOY, 1890) WOODWORD, 1906
   = Rhytidaspis murrayi McCOY, 1890 (nomen nudum)
   = Chiraropalus langtrei McCOY, 1890 (nomen nudum)
 G. primaevus EASTMAN, 1908
 G. sarlei HUSSAKOF & BRYANT, 1918
 G. parvulus BRYANT, 1929
 G? convexus GROSS, 1933
 G. rothrocki WELLS & BLICKLE, 1943
 G? eastmani WELLS, 1944
 G. warreni WHITE, 1968
 G. praecursor (ZANGERL, 1973)
   = Trichorhipsis praecursor ZANGERL, 1973
 G. seriponensis GAGNIER, TURNER, FRIMAN, SUAREZ-RIGLOS & JANVIER, 1988
 G. felius SNYDER, 2011
G. sp

Genus: Gyracanthides WOODWARD, 1906
 G. murrayi WOODWARD, 1906 (Type)
   = Gyranchthus murrayi (WOODWARD, 1906)
G. hawkinsorum emend TURNER, BURROW & WARREN, 2005
  = Gyracanthides hawkinsi TURNER, BURROW & WARREN, 2005
G. sp

Genus: Oracanthus AGASSIZ, 1837 (1843)
 O. milleri AGASSIZ, 1837 (1843) (Type)
   = Oracanthus minor AGASSIZ, 1843
   = Oracanthus confluens AGASSIZ, 1843
 O. pustulosus AGASSIZ, 1837 (1843)
 O. vetustus LEIDY, 1855 (1856)
 O? obliquus ST. JOHN & WORTHEN, 1875
 O. rectus ST. JOHN & WORTHEN, 1883
 O. bochumensis JAEKEL, 1890
 O.  lineatus NEWBERRY, 1897
 O. triangularis EASTMAN, 1917
 O. farringtoni ROY, 1929

SUBORDER: BROCHOADMONOIDEI GAGNIER & WILSON, 1996

Family: KATHEMACANTHIDAE Gagnier & Wilson, 1996

Genus: Kathemacanthus GAGNIER & WILSON, 1996
 K. rosulentus GAGNIER & WILSON, 1996 (Type)

Family: Incertae sedis

Genus: StriacanthusHILLS, 1921
 S. sicaeformis HILLS, 1921 (Type)
S. sp

ORDER: ACANTHODIFORMES BERG, 1940

Gen. sp indet.

Family: MESACANTHIDAE Moy-Thomas, 1939

Genus: Melanoacanthus CUMBAA & SCHULTZE, 2002
 M. minutus CUMBAA & SCHULTZE, 2002 (Type)

Genus: Mesacanthus TRAQUAIR, 1888
M. pusillus (AGASSIZ, 1844) TRAQUAIR, 1888
   = Acanthodes pusillus AGASSIZ, 1844
M. mitchelli (EGERTON, 1861)
   = Acanthodes mitchelli EGERTON, 1861
   = Acanthodes antiquus EGERTON, 1861
 M. peachi (EGERTON, 1861)
   = Acanthodes peachi EGERTON, 1861
   = Acanthodes coriaceus EGERTON, 1861
 M. semistriatus (WOODWARD, 1892)
   = Acanthodes semistriatus WOODWARD, 1892
M. grandis GAGNIER & GOUJET, 1997
M. sp

Genus: Promesacanthus HANKE, 2008
P. eppleri HANKE, 2008 (Type)

Genus: Teneracanthus BURROW & YOUNG, 2005
 T. toombaensis BURROW & YOUNG, 2005 (Type)
= “Gomphonchus? Sp’ TURNER et al, 1981 (fig. 14A-C).
= ‘acanthodian spines’ TURNER et al, 1981.
= ‘smloth ischnachthid scales’ YOUNG et al, 1987
= ‘climatiid spines’ TURNER, 1995.
= ‘Acanthodes-type scales’, TURNER, 1995
= ‘new mesacanthid’ YOUNG & TURNER, 2000
= ‘Acanthodian remains’ in part YOUNG & GOUJET, 2003

Genus: Triazeugacanthus MILES, 1966
 T. affinis (WHITEAVES, 1887) MILES, 1966 (Type)
   = Acanthodes affinis WHITEAVES, 1887
   = Mesacanthus affinis TRAQUAIR, 1890
   = Homalacanthus affinis RUSSELL, 1951

Family: CHEIRACANTHIDAE Berg, 1940

Genus: Carycinacanthus (ROHON, 1889) MILES, 1966
 C. lopatini (ROHON, 1889) MILES, 1966 (Type)
   = Acanthodes lopatini ROHON, 1889
= Acanthodes parvulus ROHON, 1889

Genus: Cheiracanthus AGASSIZ, 1835 (1843)
= Chiracanthus M’COY, 1846
 C. murchisoni AGASSIZ, 1835, (1843) (Type)
   = Cheiracanthus minor AGASSIZ, 1835 (1843)
   = Cheiracanthus microlepidotes AGASSIZ, 1844
   = Cheiracanthus lateralis M’COY, 1848
   = Cheiracanthus pulverulentus M’COY, 1848
 C. grandispinus M ‘COY, 1848
 C. latus EGERTON, 1861
 C. costellatus TRAQUAIR, 1893
 C. brevicostatus GROSS, 1973
 C. longiocostatus GROSS, 1973
 C. splendens GROSS, 1973
C. gibbosus VALIUKEVICIUS, 1986
 C.wangi BURROW, TURNER & WANG, 2000
C. sp

Genus: Ginkgolepis PINAKHINA & MARSS, 2018
G. tenericostatus PINAKHINA & MARSS, 2018 (Type)

Genus: Homalacanthus RUSSELL, 1951
 H. concinnus (WHITEAVES, 1887) RUSSELL, 1951 (Type)
   = Acanthodes concinnus WHITEAVES, 1887
   = Paracanthodes concinnus (WHITEAVES, 1887), GRAHM-SMITH, 1936
 H. bergi (OBRUCHEV, 1962)
   = Acanthodes bergi OBRUCHEV, 1962

Genus: Markacanthus VALIUKEVICIUS, 1985
 M. alius VALIUKEVICIUS 1985 (Type)

Family: ACANTHODIDAE Huxley, 1861
= Family: ACANTHOESSIDAE Hay, 1902

Genus: Acanthodes AGASSIZ, 1833 (1834)
= Acanthoessus AGASSIZ, 1832  (nomen nudum)
= Carycinacanthus MILES, 1966
= Rodriguezichthyes SCHULTZE, 1990
 A. bronni AGASSIZ, 1833 (1834) (Type)
   = Acanthodes rouvillei SAUVAGE, 1883
 A. sulcatus AGASSIZ, 1835
 A. gracilis (BEYRICH, 1848)
   = Holacanthodes gracilis BEYRICH, 1848
   = Holacanthodes var bendai FRITSCH, 1893 (1895)
 A. wardi EGERTON, 1866
   = Acanthodes major DAVIS, 1894
   = Acanthodes striatus WELLBURN, 1901
 A. pristis CLARKE, 1885 (nomen dubium)
A. lopatini ROHON, 1889
= Homalacanthus lopatini (ROHON, 1889) NOVITSKAYA & OBRUCHEV, 1964
= Carycinacanthus lopatini (ROHON, 1889) MILES, 1966
= Acanthodes parvulus ROHON, 1889
= Acanthodes bergi OBRUCHEV, 1962
= Homalacanthus bergi (OBRUCHEV, 1962)
 A. nitdus WOODWARD, 1891
 A. punctatus FRITSCH, 1893 (1895)
 A. beecheri EASTMAN, 1902
 A. marshi EASTMAN, 1902
 A? dubliensis STAUFFER, 1938 (nomen dubium)
   = Acanthoides hardyi WELLS, 1944
   = Acanthoides sciotoensis WELLS, 1944
 A. australis WOODWARD, 1906
 A. ovensi WHITE, 1927
 A. bridgei ZIDEK, 1976
 A. lundi ZIDEK, 1980
 A. luedersensis (DALQUEST, et al, 1988)
   = Rodriguezichthyes luedersensis SCHULTZE, 1990
A. guizhouensis WANG & TURNER, 1985
 A. tholeyi HEIDTKE, 1990
 A. kinneyi ZIDEK, 1992
A. sp

Genus: Eupleurogmus McCOY, 1890
 E. creswelli McCOY, 1890 (Type)

Genus: Howittacanthus LONG, 1986b
 H. kentoni LONG, 1986b (Type)

Genus: Paracanthodes

Genus: Protogonacanthus MILES, 1966
 P. juergeni MILES, 1966 (Type)

Genus: Pseudacanthodes WHITE & MOY-THOMAS, 1941
 P. pinnatus (FRITSCH, 1893) WHITE & MOY-THOMAS, 1941 (Type)
   = Protacanthodes pinnatus FRISCH, 1893

Genus: Traquairichthys WHITLEY, 1933
= por Traquarira FRITSCH, 1893 (1895) non CARRUTHERS, 1872
 T. pygmaeus (FRITSCH, 1875) WHITLEY, 1933 (Type)
   = Acanthodes pygmaeus FRITISCH, 1875
T. sp

Genus: Utahacanthus SCHULTZE, 1990
 U. gutheri SCHUTLZE, 1990 (Type)

Genus: Westrichus HEIDTKE, 2003
 W. kraetschmeri HEIDTKE, 2003 (Type)

Gen. sp indet.

_____________________________________________________________________________________


Bernacsek, G. M., and Dineley, D. L., 1977, New acanthodians from the Delorme Formation (Lower Devonian) of N.W.T., Canada: Palaeontographica Abt A, v. 158, lfg. 1-3, p. 1-15.

Blais, S. A., Hermus, C. R., and Wilson, M. V. H., 2015, Four new early Devonian Ischnacanthid acanthodians from the Mackenzie Mountains, Northwest Territories, Canada: An early experiment in dental diversity: Journal of Vertebrate Paleontology, v. 35, n. 1, e948546, 13pp.

Bryant, W. L., 1934, The fish fauna of Beartooth Butte, Wyoming, Parts II and III: Poceedings of the American Philosophical Society, v. 73, p. 127-162.

Burroy, C. J., Janvier, P., an dVillarroel, C., 2003, Late Devonian acanthodians from Colombia: Journal of South American Earth Sciences, v. 16, p. 155-161.

Burrow, C. J., 2002, Lower Devonian acanthodian fuans and biosratigrpahy of south-eatern Australia: Memoirs of the Association of Australasian Palaeontologist, v. 27, p. 75-137.

Burrow, C. J., Janvier, P., and Villarroel, C., 2003, Late Devonian acanthodians from Colombia: Journal of South American Earth Sciences, v. 16, p. 155-161.

Burrow, C. J., Lelievre, H., and Janjou, D., 2006, Gnathostome microremains from the Lower Devonian Jawf Formation, Saudi Arabia: Journal of Paleontology, v. 80, n. 3, p. 537-560.

Burrow, C. J., Long, J. A., and Trinajstic, K., 2009, Disarticulated acanthodian and chondrichthyan remains from the upper Middle Devonian Aztec Siltstone, southern Victoria Land, Antarctica: Antarctic Science, v. 21, n. 1, p. 71-88.

Burrow, C. J., and Rudkin, D., 2014, Oldest near-complete acanthodian: the first vertebrate from the Silurian Bertie Formation Konservat-Lagerstatte, Ontario: Public Library of Science (PLOS), One, v. 9, n. 8, 7 pp.

Burrow, C. J., Vergoossen, J. M. J.,Turner, S., Uyeno, T. T., and Thorsteinsson, R., 1999, Microvertebrate assemblages from the Upper Silurian of Cornwallis Island, Arctic, Canada: Canadian Journal of Earth Science, v. 36, p. 349-361.

Burrow, C. J., and Young, G. C., 1999, An articulated teleostome fish from the Late Silurian (Ludlow) of Victoria, Australia: In: Papers in Vertebrate paleontology, edited by Baynes, A., and Long, J. A., Records of the Western Australian Museum, Supplement n. 57, p. 1-14.

Burrow, C. J., and Young, G. C., 2005, The acanthodian fauna of the Craven Peaks Beds (Early to Middle Devonian), Western Queensland: Memoirs of the Queensland Museum, v .51, n. 1, p. 3-25.

 

Cumba, S. L., and Schultz, H.-P., 2002, An Early Devonian (Emsian) acanthodian from the Bear Rock Formation, Anderson River, Northwest Territories, Canada: Canadian Journal of Earth Sciences, v. 39, p. 1457-1465.

 

Eastman, C. R., 1907, Devonic fishes of the New York Formations: New York State Museum, memoir 10, 235pp.

Eastman, C. R., 1908, Devonian fishes of Iowa: Iowa Geological Survey, v. 18, p. 29-386.

 

Forey, P. L., Young, V. T., and McClure, H. A., 1992, Lower Devonian fishes from Saudi Arabia. Bull. Bulletin of the Natural History Museum of London, (Geology series), v. 48, n. 2, p. 25-43.

 

Gagnier, P.-Y., and Goujet, D., 1997, Nouveaux pooissons acanthodiesn du Devonien du Spitsberg: Geodiversitas, v. 19, n. 3, p. 505-513.

Gagnier, P.-Y., Turner, S., Friman, L., Suarex-Riglos, M., and Janvier, P., 1988, The Devonian vertebrate and mollusc fauna from Seripona (Dept. Of Chuquisaca, Bolivia). Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, v. 176, n. 2, p. 269-297.

Gagnier, P.-Y., and Wilson, M. V. H., 1996, Early Devonian Acanthodians from Northern Canada: Palaeontology, v. 39, part 2, p. 241-258.

Gess, R. W., 2001, A new species of Diplacanthus from the Late Devonian (Famennian) of South Africa: Annales of Paleontology, v. 87, n. 1, p. 49-60.

Groß, W., 1971. Downtonische und Dittonische Acanthodier-reste des ostseegebietes: Palaeontographica, Abt. A., v. 136, lfg. 1-6, p. 1-82.

Groß, W., 1973, Kleinschuppen, flossenstachlen und zahne von fischen aus Europaischen und Nordamerikanischen bonebeds des Devons: Palaeontographica Abt. A., v. 142, Lfg. 1-6, p. 51-155.

 

Hairapetian, V,. Valiukevicius, J, and Burrow, C. J., 2006, Early Frasnian acanthodians from Central Iran: Acta Palaeontologica Polonica, v. 51, n. 2, p. 499-520.

Heidtke, U., 1990, Studien uber Acanthodes. 3. Acanthodes tholeyi n. sp., ein neuer Acanthodier (Acanthodii: Pisces) aus dem Rotliegenden (Unterperm) der Saar-Nahe-Senke (Sudwest-Deutschland): Paläontologische Zeitschrift, v. 64, n. ¾, p. 261-267.

Hanke, G. F., 2002, Paucicanthus vanelsti gen. et sp. nov., an Early Devonian Lochkovian) acanthodian that lacks paired fin-spines: Canadian Journal of Earth Science, v. 39, p. 1071-1083.

Hanke, G. F., 2008, Promesacanthus eppleri n. gen., n. sp., a mesacanthid (Acanthodii, Acanthodiformes) from the Lower Devonian of northern Canada: Geodiversitas, v. 30, n. 2, p. 287-302.

Hanke, G. F., Wilson, M. V. H., and Lindoe, L. A., 2001, New species of Silurian acanthodians from the Mackenzie Mountains, Canada: Canadian Journal of Earth Science, v. 28, p. 1517-1529.

Heidtke, U., 2003, Westrichus kraetschmeri n. g. n. sp., ein weiterer Acanthodier (Pisces: Acanthodii) aus dem Unteren Rotliegend des Saar-Nahe-Beckens (Karbon-Perm-Grenzberich: SW-Deutschland): Neues Jahrbuch für Geologie und Paläontologie Monatschefte, heft 5, p. 257-282.

Hussakof, L., and Byrant, W. L., 1918, Catalog of the Fossil Fish in the Museum of the Buffalo Society of Natural Sciences: Bulletin of the Buffalo Society of Natural Sciences, v. 12, p. 1-343.

 

Karatajute-Talimaa, V., and Smith, M. M., 2003 (for 2002), Early acanthodians from the Lower Silurian of Asia: Transactions of the Royal Society of Edinburgh: Earth Sciences, v. 93, p. 277-299.

 

Li, Q., Cui, X., Andreev, P. S., Zhao, W., Wang, J., Peng, L., and Zhu, M., 2021, Nostolepis scale remains (Stem Chondrichthyes) from the Lower Devonian of Qujing, Yunnan, China: PeerJ 9:e11093 http://doi.org/10.7717/peerj.11093, 21pp.

Lindley, L. D., 2000, Acanthodian fish remains from the Lower Devonian Cavan Bluff Limestone (Murrumbidgee Group), Taemas district, New South Wales: Alcheringa, v. 24, p. 11-35.

Long, J. A., 1986a, New ischnacanthid acanthodians from the Early Devonian of Australia, with comments on acanthodian interrelationships: Zoological Journal of the Linnean Society, v. 87, p. 321-339.

Long, J. A., 1986b, A new late Devonian Acanthodian fish from Mt. Howitt, Victoria, Australia, with remarks on Acanthodian biogeography: Proceedings of the Royal Society of Victoria, v. 98, n. 1, p. 1-17.

Long, J. A., Burrow, C. J., and Ritchie, A., 2004, A new Late Devonian acanthodian fish from the Hunter Formation near Grenfell, New South Wales: Alcheringa, v. 28, p. 147-156.

 

Miles, R. S., 1973, Articulated Acanthodian Fishes from the Old Red Sandstone of England, with a Review of the structure and Evolution of the Acanthodian Shoulder-girdle: Bulletin of the British Museum (Natural History) Geology, v. 24, n. 2, p. 113-213.

Miller, C. G., and Marss, T., 1999, A conodont, thelodont and acanthodian fauna from the Lower Pridoli (Silurian) of the much Wenlock Area, Shropshire: Palaeontology, v. 42, part 4, p. 691-784.

 

Newberry, J. S., 1873, Descriptions of fossil fishes: Geological Survey of Ohio, Paleontology, v. 1, part II, p. 247-355.

Newman, M. J., Burrow, C. J., and den Blaauwem, J. L., 2020, A new species of ischnacanthiform acanthodian from the Givetian of Mimerdalen, Svalbard: Norwegian Journal of Geology, v. 99, n. 4, p. 1-13.

 

Orvig, T., 1967, Some new acanthodian material from the Lower Devonian of Europe: In: Papers on Fossil Vertebrates, edited by Patterson, C., and Greenwood, P. H., The Journal of the Linnean Society of London (Zoology), v. 47, n. 311, p. 131-153.

P’an, K., 1964, Some Devonian and Carboniferous fishes from South China: Acta Palaontologica Sincia, v. 12, n. 1, p. 139-168.
Pinakhina, D. V., and Marss, T., 2018, The Middle Devonian acanthodian assemblage of the Karksi outcrop in Estonia: Estonia Journal of Earth Sciences, v. 67, n. 1, p. 96-111.

 

Reed, J. W., 1986, The Acanthodian genera Machaeracanthus and Persacanthus from the Devonian of Red Hill, Nevada: Geobios, n. 19, fasc. 4, p. 409-419.

 

Schultze, H.-P., 1982, Ein primitiver Acanthodier (Pisces) aus dem  Unterdevon Lettlands: Paläontologische Zeitschrift, v. 56, n. ½, p. 95-105.

Schultz, H.-P., 1990, A new acanthodian from the Pennsylvanian of Utah, U.S.A., and the distribution of otoliths in gnathostomes: Journal of Vertebrate Paleontology, v. 10, n. 1, p. 49-58.

Synder, D., 2011, Gyracanthid gnathostome remains from the Carboniferous of Illinois: Journal of Vertebrate Paleontology, v. 31, n. 4, p. 902-906.

Südkamp, W. H., Burrow, B., and Burrow, C. J., 2007, The acanthodian Machaeracanthus from the Lower Devonian Hunsrück Slate of the Hunsrück region (Germany): Paläontologische Zeitschrift, v. 81, Heft 1, p. 97-104.

 

Traquair, R. H., 1894, Notes on Palaeozoic fishes: Annals & Magazine of Natural History, series 6, v. 14, p. 368-374.

Turner, S., Burrow, C. J., and Warren, A., 2005, Gyracanthides hawkinsi sp. nov. (Acanthodii, Gyracanthidae) from the Lower Carboniferous of Queensland, Australia, with a review of Gyracanthid taxa: Paleontology, v. 48, part 5, p. 963-1006.

Valiukevicius, J., 1992, First articualted Poracanthodes from the Lower Devonian of Severnaya Zemlya: In: Fossil Fishes as Living Animals. Academia 1,edited by Mark-Kurik, E., Institue of Geology, p. 193-213.

Valiukevicius, J., 1998, Acanthodians and zonal stratigraphy of Lower and Middle Devonian in East Baltic and Byelorussia: Palaeontographica Abt. A., v. 248, Lfg. 1-2, p. 1-53.

Valiukevicius, J., 2003, Devonian acanthodians from Severnaya Zemlya Archipelago (Russia): Geodiversitas, v. 25, n. 1, p. 131-204.

Valiukevicius, J., 2004, New Wenlock – Pridoli (Silurian) acanthodian fishes from Lithuania: Acta Palaeontologica Polonica, v. 49, n. 1, p. 147-160.

Valiukevicius, J., 2004, Silurian acanthodian succession of the Luzni-4 borehole (Latvia): In: The Second Gross Symposium "Advances of Palaeoichthyology", Earth and Environment Sciences, Acta Universitatis Latviensis, v. 679, p. 120-147.

Valiukevicius, J., and Burrow, C. J., 2005, Diversity in tissues in acanthodians with Nostolepis-type histological structure: Acta Palaeontologica Polonica, v. 50, n. 3, p. 635-649.

Vergossen, J. M. J., 1999, Late Silurian fish microfossils from Helvetesgraven, Skane (southern Sweden) (I)): In: Proceedings of the Third European Workshop on Vertebrate Paleontology, (Maastricht, May 6-9, 1998). Geology Mijnbouw, v. 78, n. 3-4, p. 267-280.

Vergoossen, J. M. J., 1999, Late Siluran fish microfossils form an East Baltic-derived erratic from Oosterhaule, with a description of a new acanthodian taxa: Geolgoie en Mijnbouw, v. 78, p. 231-251.

Vidal, M., Goujet, D., Janier, P., and Lardeuz, H., 1994, Micro-restes de Vertebres du Calciare de La Grange, Devonien inferieur (Emsien) du Sud-Est du Massif Armoricain: Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, v. 194, n.  2/3, p. 321-341.

Vieth-Schreiner, J., 1983, Fische-schuppen und zahne aus der eifeler Kalkmulden Zone (Emisum, Eifelium): Senckenbergiana Lethaea, v. 64, n. 2/4, p. 129-177.

Voichyshyn, V., and Szanianwski, 2012, Acanthodiani jaw bones from Lower Devonian marine deposits of Podolia, Ukraine: Acta Palaeontologica Polonica, v. 57, n. 4, p. 879-896.

Voichyshyn, V., and Szaniawski, H., 2018, New Ischnacanthiform jaw bones from the Lower Devonian of Podolia, ukraine: Acta Palaeontologica Polonica, v. 63, n. 2, p. 327-339. \

Wang, N.-Z., 1984, Thelodont, Acanthodian, and Chondrichthyan fossils from the Lower Devonian of southwest China: In: Evolution & Biogeography of Early Vertebraets, Proceedings of the Linnean Socity of New South Wales, v. 107, n. 3, p. 419-441.

Wang, N.-Z., Wang, J., Zhang, G., and Wang, S., 1998, The first discovery of Silurian and Early Devonian Acanthoians from Zoige and Tewo Counties, West Qinling Mountains: Vertebrata Pal Asiatica, v. 36, n. 4, p. 268-281.

Wang, S., and Turner, S., 1985, Vertebrate microfissils of the Devonian-Carboniferous Boundary, Muhua section, Guizhou Province: Vertebrata PalAsiatica, v. 23, n. 3, p. 223-234.

Wang, W., 2003, First occurrence of acanthodian microfossils from the Early Devonian of Lijiang, Yunnan, China: Vertebrata PalAsiatica, v. 41,n. 1, p. 1-16.

White, E. I., 1926, The fish-fauna of the Cementstones of Foulden, Berwickshire: Transcations o fthe Royal Society of Edinburgh, v. 55, part 1, p. 11, p. 255-286.

 

Young, G. C., 1989, The Aztec fish fauna (Devonian) of Southern Victoria Land: Evolutionary and biogeographic significance: In: Origins and Evolution of the Antartic Biota, editor Crame, J. A., Geological Society Special Publication, n. 47, p. 43-62.

Young, G. C., 1989, New occurrences of Culmacanthid Acanthodians (Pisces, Devonian) from Antarctica and southeastern Australia: Proceedings of the Linnean Society of New South Wales, v. 111, n. 1, p. 11-24.

 

Zidek, J., 1976, Kansas Hamilton Quarry (Upper Pennsylvanian) Acanthodes, with remarks on the Previously reported North American occurrences of the Genus: The University of Kansas Paleontological Contributions, Paper 83, p. 1-41.

Zidek, J., 1980, Acanthodes lundi, a new species (Acanthodii) and associated coprolites from the Uppermost Mississippian Heath Formation of Central Montana: Annals of Carnegie Museum, v. 49, article 3, p. 49-78.